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However, we estimate the crown age of the five orchid subfamilies to be generally (~1-8 Ma) younger than previously calculated under the Penalized Likelihood algorithm and using a single internal fossil calibration.The crown age of Contrary to recent suggestions that orchid diversification began in a period of global warming, our results place the onset of diversification of the largest orchid subfamilies (Orchidoideae and Epidendroideae) in a period of global cooling subsequent to the Early Eocene Climatic Optimum.BEAST includes software libraries from projects whose licensing terms require that the following notices be included when they are distributed in binary form and that the recipient be given a copy of their respective licenses.· This product includes software developed by the JDOM Project ( · This product includes software developed by The Apache Software Foundation ( · The license for JDOM can be read in the file in the source distribution available at · The license for the Commons Math library can be read at or in the file in the source distribution available at This manual page and its XML source can be used, modified, and redistributed as if it were in public domain.For us it simply means that the interface will be the same on all platforms.The screenshots used in this tutorial are taken on a Mac OS X computer; however, both programs will have the same layout and functionality on both Windows and Linux.

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These areas are species phylogenies for molecular dating, coalescent-based population genetics and measurably evolving populations (ancient DNA or time-stamped viral sequence data sets).· Drummond AJ, Rambaut A & Shapiro B and Pybus OG (2005) Mol Biol Evol 22, 1185-1192. Suchard (BEAST) Copyright © 2009–2010 Felix Feyertag (This manpage) The binaries and source code are made available and can be distributed subject to the following conditions: BEAST is Copyright (C) 2002-2006 Alexei Drummond and Andrew Rambaut BEAST is distributed under the terms of the GNU Lesser General Public License.· Drummond AJ, Nicholls GK, Rodrigo AG & Solomon W (2002) Genetics 161, 1307-1320. See the notice in individual files and the file LICENSE for licensing details.The temporal origin and diversification of orchids (family Orchidaceae) has been subject to intense debate in the last decade.The description of the first reliable fossil in 2007 enabled a direct calibration of the orchid phylogeny, but little attention has been paid to the potential influence of dating methodology in obtaining reliable age estimates.

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